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 External Resources

 48138https://www.addgene.org/48138  Addgene identifier (48138, reused)

 9606https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?id=9606  NCBI taxonomy (human, Homo sapiens)

 0000-0003-2917-0778https://orcid.org/0000-0003-2917-0778  ORCID identifier (Steffen J. Sahl)

 0000-0001-6560-2174https://orcid.org/0000-0001-6560-2174  ORCID identifier (Jessica Matthias)

 0000-0001-5959-6409https://orcid.org/0000-0001-5959-6409  ORCID identifier (Kaushik Inamdar)

 0000-0001-9680-7543https://orcid.org/0000-0001-9680-7543  ORCID identifier (Michael Weber)

 0000-0001-5483-5837https://orcid.org/0000-0001-5483-5837  ORCID identifier (Taukeer A. Khan)

 0000-0003-1873-5120https://orcid.org/0000-0003-1873-5120  ORCID identifier (Christian Brüser)

 0000-0002-8028-3121https://orcid.org/0000-0002-8028-3121  ORCID identifier (Stefan Jakobs)

 0000-0003-2041-5740https://orcid.org/0000-0003-2041-5740  ORCID identifier (Stefan Becker)

 0000-0002-1266-4344https://orcid.org/0000-0002-1266-4344  ORCID identifier (Christian Griesinger)

 0000-0003-3084-6595https://orcid.org/0000-0003-3084-6595  ORCID identifier (Johannes Broichhagen)

 0000-0002-9638-5077https://orcid.org/0000-0002-9638-5077  ORCID identifier (Stefan Walter Hell)

 1/518https://mbexc.uni-goettingen.de/archive/1/518  Research Data Archive (supplemental material)

 1IGYhttps://www.rcsb.org/structure/1IGY  RCSB PDB Identifier (DOI:10.2210/pdb1IGY/pdb, reused)

 1IVThttps://www.rcsb.org/structure/1IVT  RCSB PDB Identifier (DOI:10.2210/pdb1IVT/pdb, reused)

 4HZGhttps://www.rcsb.org/structure/4HZG  RCSB PDB Identifier (DOI:10.2210/pdb4HZG/pdb, reused)

 5FQ1https://www.rcsb.org/structure/5FQ1  RCSB PDB Identifier (DOI:10.2210/pdb5FQ1/pdb, reused)

 6I2Ghttps://www.rcsb.org/structure/6I2G  RCSB PDB Identifier (DOI:10.2210/pdb6I2G/pdb, reused)

 01s1h3j07https://ror.org/01s1h3j07  ROR identifier (01s1h3j07, Fraunhofer Institute for Translational Medicine and Pharmacology)

 03av75f26https://ror.org/03av75f26  ROR identifier (03av75f26, Max Planck Institute for Multidisciplinary Sciences)

 000bxzc63https://ror.org/000bxzc63  ROR identifier (000bxzc63, Max Planck Institute for Medical Research)

 05xy1nn52https://ror.org/05xy1nn52  ROR identifier (05xy1nn52, Multiscale Bioimaging)

 021ft0n22https://ror.org/021ft0n22  ROR identifier (021ft0n22, University Medical Center Göttingen)

 CVCL_0042https://scicrunch.org/resolver/CVCL_0042  SciCrunch identifier (RRID:CVCL_0042, U2OS)

 SCR_016387https://scicrunch.org/resolver/SCR_016387  SciCrunch identifier (RRID:SCR_016387, Illumina NovaSeq 6000 Sequencing System)

 SCR_001622https://scicrunch.org/resolver/SCR_001622  SciCrunch identifier (RRID:SCR_001622, MATLAB)

 science....sm.pdfhttps://www.science.org/doi/suppl/10.1126/science.adj7368/suppl_file/science.adj7368_sm.pdf  Supplemental material

 science....st.pdfhttps://www.science.org/doi/suppl/10.1126/science.adj7368/suppl_file/science.adj7368_mdar_reproducibility_checklist.pdf  Supplemental material (MDAR reproducibility checklist)

 mbexc.de...ision/https://mbexc.de/direct-optical-measuerement-of-intramolecular-distances-with-angstrom-precision/  Website entry (mbexc.de)

 10.5281/zenodo.12744422https://doi.org/10.5281/zenodo.12744422  Zenodo identifier (10.5281/zenodo.12744422)